See Robin's paper on this:
http://hanson.gmu.edu/hardstep.pdf
If a step is extremely hard (and thus astronomically unlikely to occur in the lifespan of a planet) then we should expect to see it taking a length of time comparable to typical planetary lifespans divided by the number of steps.
The last two steps occurred suspiciously quickly to be super-hard, and the independent clusters of mammalian braininess in cetaceans and primates [EDIT: (and birds, to a lesser extent)] make the third step questionable too.
Super-difficult life and nervous systems look very plausible to me. A special difficulty involved in the formation of mammals (a feature that predisposed to the later development of intelligent hominids) seems less plausible but not very implausible.
In addition to steps that are hard in the sense that they can take a long time, there may also be tricky steps -- ones that have a finite window of opportunity before being precluded by some other version of events. I don't have a good enough theory of the development of civilization to defend any candidates in history, but here's a paleontological candidate:
Consider that animal lineages readily lose limbs but, once lost, almost never regain them (many-legged arthropods to six-legged insects, four-legged reptiles to two-legged birds and no-legged snakes, but no reverse transitions; the reasons for this are easy to understand in terms of the requirement of evolution that intermediate forms be advantageous). It is also clear that animals can get by well enough with two legs, but sparing two of four for toolmaking has always been a problem, let alone sparing two of two. Finally, it is clear that four legs over two is not necessarily enough advantage to displace an entrenched competitor.
The four-legged lungfish that became the ancestor of amphibians could not of course have known its distant descendents would need two spare limbs for toolmaking. If it had been delayed until a two-legged lungfish had taken the niche... it is not certain that the ultimate development of civilization would have been rendered impossible, but it is at least plausible.
One question we need to ask about the question of time is what sort of process leads to each breakthrough.
Is it more like buying a lottery ticket with every generation, or is it like saving until you have enough money to buy the next step?
It may well be that the Cambrian explosion was the result of 3 billion years of small improvements and would have been impossible at 1 billion years.
The history of invention in human history seems to work more like savings -- as soon as sufficient progress has been made, the breakthrough happens independently.
There's a problem in independent invention in evolution as well--once the first evolution takes place that niche is occupied. An independent invention may be beat out for resources by the more polished first-mover. Short-lived species leave very few fossils.
Something at least remotely analogous to the nervous system seems to have evolved in some plants, which use cells with something like action potentials to drive rapid movement or rapid changes of chemical behavior in response to environmental stresses.
Bats also have high encephalization quotients, and corvids seem to have the sorts of behaviors characteristic of high encephalization quotients, though both may be too absolutely small to become civilized absent unusual environmental conditions.
The latter three items all seem sufficiently rapid to provide l...
There were wheels in the western hemisphere before the arrival of the europeans, but they were only used for toys. No one seemed to guess that they might be useful. But if they'd had more time....
ETA: perhaps I should cite Diamond here in case anyone wonders where this factoid comes from. It's from "Guns, Germs, and Steel".
Multicellularity seems to have evolved multiple times independently
This isn't really true. Only organisms with mitochondria developed multicellularity. Mitochondria are the hard part.
eusociality developed in aphids, thrips, mole rats, termites, and at least 11 times in Hymenoptera
Similarly, it would be more informative to say that Hymenoptera developed a particular pattern of chromosomal inheritance once, and that led to 11 different types of eusocialism.
Re: abiogenesis. You say:
we know of no mechanism under which creation of life seems even remotely plausible.
For a plausible mechanism, see this video. (It starts with anti-creationism stuff; skip to 2:45 to watch the science.)
Our observations are biased because anything that occurs multiple times is very easy to see but something that occurs only once could be completely missed as an essential step towards civilization because we assume it was inevitable.
AK's Rambling Thoughts has an interesting post on the originas of bilaterians (as well as eukaryotes and life in general).
Nervous system evolved only once, about 3 billion years after life started, and nothing analogous to it ever evolved in any other lineage. [Urbilaterian]
From Science, July 3 2009, p. 24-26, "On the origin of the nervous system":
...Assembling these components into a cell a modern neuroscientist would recognize as a neuron probably happened very early in animal evolution, more than 600 million years ago... Scientists also disagree on which animals were the first to have a centralized nervous system and how many times neurons and nervous sytems ev
Life did not die out on Earth, or on any particular environment where it previously thrived, in spite of major changes in temperature, composition of atmosphere, and multiple large scale disasters. This suggests life is very resilient. Every time life is wiped out in some part of Earth, it is quickly recolonized.
Be careful of anthropic bias here. Taken alone, the argument "life did not die out on Earth" is invalid because if it had, we wouldn't be here. However, the second point, that when some evolutionary niche is wiped out it is quickly col...
One thing that caught my eye is the presentation of "Universe is not filled with technical civilizations..." as data against the hypothesis of modern civilizations being probable.
It occurs to me that this could mean any of three things, which only one of which indicates that modern civilizations are improbable.
1) Modern civilizations are in fact as rare as they appear to be because they are unlikely to emerge. This is the interpretation used by this article.
2) Modern civilizations collapse quickly back to a premodern state, either by fighting a v...
One other thing an advanced technological civilization seems to need is concentrated energy. We're highly dependent on coal and oil. At this stage, nuclear could be substituted, but I don't know that there would have been enough slack for the research to get nuclear without the fossil fuels.
It seems plausible that any planet which has had extensive life for long enough to develop intelligence would also have fossil fuels, but that's pretty vague. It doesn't guarantee that the fossils don't get dispersed, eaten, or end up too deep to be easily accessible.
I'...
No we're not. The data is clearly against this theory.
Coal was barely used until 1800s, early industrial revolution machinery used wood (indirectly solar power), charcoal, and river flow (indirectly solar power) instead. Oil didn't matter much until 1950s.
Amount of solar energy Earth receives annually is 3,850,000 EJ (and if we ever needed more there are ridiculously higher amounts of solar energy available is space). Human primary energy use is 487 EJ, or 0.01% of that. That's of course only because we conveniently don't count solar energy used to grow our food, and heat our planet - otherwise it would be fair to say human civilization uses 99.99% solar power (via photosynthesis, heating, water flow, wind etc.) and 0.01% all the other kinds of energy like fossil fuels, nuclear, geothermal etc.
We know fossil fuels were not necessary for industrial civilization because by the time we started using them we were already had industrial civilization. That's as good a proof as it gets.
History of ferrous metallurgy History of coal mining History of petroleum Solar energy
EDIT: Also, long before railways, river transport, and long distance sea transport were extremely common. If some place ...
Primates existed for about 85mln years, Cetaceans for about 48mln years, in all this time nothing got even close to Homo except for Homo. (Evolution of cetaceans Evolution of primates) Of course plenty of other large social animals had an opportunity to develop something like Homo for at least 300mln years before that. (Evolution of Reptiles), or more if we seriously treat possibility of intelligent life in the sea.
So life, animals with nervous system (these are linked, as without nervous system animal complexity is very small), and Homo seem safe.
The case for language and behavioral modernity seems weaker indeed. One obvious argument is that Robin's paper doesn't allow reversal to previous states (die-off of genus Homo - something very likely), so if expected lifespan of Homo genus (due to all ecological changes, and assuming no civilization - the bottleneck event theory suggests it's not too unlikely) was 10mln years, and we expect something like Homo to appear once, then if it took 3mln years, it's 30% of its possible time, what sounds reasonably hard.
Another case for language is that there is no obvious mechanism how it could have evolved, so perhaps our Homo was luckily predisposed for it. And there's no reason why you need Homo for something like language - if chimps or dolphins could be taught grammar, that would be evidence for language being easy - yet it seems that in 600mln years of animals with brains nothing like it have happened.
Language and behavioral modernity are so close that they might be extremely closely related. If they're independent (assuming Homo), we can use expected genus extinction argument.
Or if language turns out to be far older, happening together with Homo (seems unlikely but not impossible), then behavioral modernity needs explanation instead.
Timing as we know it is:
With exact timing of development of language being unknown.
Good point about the extinction of lineages, TAW. Updating.
We have a sample of one modern human civilization, but there are some hints on how likely it was to happen.
Major types of hints are:
Data for:
Data against:
To me it looks like life, animals with nervous systems, Upper Paleolithic-style Homo, language, and behavioral modernity were all extremely unlikely events (notice how far ago they are - vaguely ~3.5bln, ~600mln, ~3mln, ~200k or ~600k, ~50k years ago) - except perhaps language and behavioral modernity might have been linked with each other, if language was relatively late (Homo sapiens only) and behavioral modernity more gradual (and its apparent suddenness is an artifact). Once we have behavioral modernity, modern civilization seems almost inevitable. Your interpretation might vary of course, but at least now you have a lot of data to argue for your position, in convenient format.