Jowibou comments on Attention Lurkers: Please say hi - Less Wrong
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Also, the karma system adds an additional barrier, at least in my mind. Knowing that your comment is going to be explicitly judged and your score added to a "permanent record" can be intimidating.
Whether we like it or not, that "intimidation" may be the single most important factor in keeping the level of discourse in the comments unusually high. Status games can be beneficial.
Indeed. I'm not saying the karma system is a bad thing.