zero_call comments on You're Entitled to Arguments, But Not (That Particular) Proof - Less Wrong
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Probabilities can't be ignored, of course, but nobody ever actually has the correct probabilities, except for in mathematics and experimental science. (Although in mathematics, you might say the correctness is only platonic.) When you say "probabilities" in this article, IMO, you are rather blase about interchanging the notions of these objective, accurate entities, and these nonobjective entities that get mixed up alongside of them.
The whole point of making an objective claim should be to attempt to provide evidence for why your probabilities are accurate. Thus the initial quote you mentioned about the burden of evidence is actually completely crucial and well justified, in my mind. In a situation where you can't establish this accuracy because evidence is lacking, there's really no other resort except to be skeptical, or to rely on the opinion of experts who are better able to interpret more difficult evidence.
The problem with the evolution example is that the theistic person isn't very well characterized by their skepticism. On the contrary, I would say they are more characterized by their willingness to believe, and they are a reverse-skeptic, who is unswayed by evidence but also unswayed by lack of evidence. I see much more of a problem with this kind of attitude than with the attitude of "over-skepticism" (to crudely paraphrase) that you've alluded to here.
Probability is a strength of belief. Even if you are not able to calculate that strength precisely, that strength should respond to evidence in a certain way, and there is a rational probability to assign a proposition given the evidence observed.
The problem with the theist is not being consistently over or under skeptical, but being more skeptical about propositions they don't want to believe, so they are over-skeptical of evolution in the way that Eliezer describes.