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Barry_Cotter comments on Forager Anthropology - Less Wrong
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Nobody can downvote on Hacker News. The only vaguely analogous function is "flag" which leads to posts (not comments) being killed or marked for killing.
(Edit:rhollerithdotcom points out correctly that this is only true for submissions and that above a karma threshold comments are downvotable)
Useful essay on online communities http://www.kuro5hin.org/story/2009/3/12/33338/3000
If one has enough karma (ISTR the threshhold being 200 points at one point, though it has probably been raised a few times since then) one can downvote comments or else how to explain the presence of comments with negative scores in almost every comment section.
You might be right about top-level submissions though.