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Barry_Cotter comments on Forager Anthropology - Less Wrong
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How much experience have you had watching the trajectory of online communities?
Have you for example informed yourself of the case of Reddit (the original one) which is particularly relevant to this community in that the software is so similar?
I have not, but Paul Graham has (since he was an investor in Reddit) and he has stated many times that he believes that his community, Hacker News, is in constant danger of falling prey to the dynamic that rendered Reddit worthless to thoughtful busy people, and he has taken many different measures, including banning a user relatively frequently, denying new users the right to cast downvotes -- or any votes at all if their karma is low enough -- and disappearing the "reply" link on certain posts based on an algorithm.
Nobody can downvote on Hacker News. The only vaguely analogous function is "flag" which leads to posts (not comments) being killed or marked for killing.
(Edit:rhollerithdotcom points out correctly that this is only true for submissions and that above a karma threshold comments are downvotable)
Useful essay on online communities http://www.kuro5hin.org/story/2009/3/12/33338/3000
If one has enough karma (ISTR the threshhold being 200 points at one point, though it has probably been raised a few times since then) one can downvote comments or else how to explain the presence of comments with negative scores in almost every comment section.
You might be right about top-level submissions though.