As Wright says, organisms are not that harmonious anyway. At every polymorphic locus, two alleles are engaged in a battle-to-the-death.
There is still enough cooperation for people to describe the resulting behaviour as "purposeful" - despite all the underlying replicator-level conflicts that produce it.
Sweet, there's another Bloggingheads episode with Eliezer.
Bloggingheads: Robert Wright and Eliezer Yudkowsky: Science Saturday: Purposes and Futures