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ata comments on Transhumanism and the denotation-connotation gap - Less Wrong
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I figured it was just a joke playing on the common assertion that something is what "makes us human", something discussed (and, to a certain extent, deconstructed) in the post.
A joke, and an example of using a more-precise word instead of the most-accessible word. Other mammals don't speak, but I'd bet they have concepts and contexts activated by those concepts. Invertebrates seem to have less-flexible, less context-sensitive categories; so my guess as to what having connotations distinguishes us from, is invertebrates.
That does seem far more likely :)