timtyler comments on Group selection update - Less Wrong
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That seems to be a rather confused way of putting it. Or course selection operates between species. The issue is whether it results in very much in the way of species-level adaptations.
Did you read the whole abstract? They say "species selection is an important process":
I did read the whole abstract: the author admits that species selection is controversial in modern evolutionary biology, and in the rest of the paper argues that this should not be the case. The point of my previous comment was not whether species selection should or should not be recognized as important, because I do not know. It was a question concerning how well-accepted species selection is amongst biologists.