kilobug comments on On Being Okay with the Truth - Less Wrong
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I would say that "becoming strong and opressing the weak" is the default goal. You don't need any kind of morality here, it's just biology of a social species. Being strong has natural rewards.
Morality is what allows you to have alternative goals. Morality means that "X is important too", sometimes even more important than being strong (though usually it is good to both be strong and do X). Morality gives you social rewards for doing X.
Being strong is favored by genes, doing X is favored by (X-promoting) memes. In the absence of memes (more precisely in absence of strong memes saying what is right and wrong), humans fall back on their natural social behavior, the pecking order. In the presence of such memes, humans try to do X; and also at the same time secretly try to be strong, but they cannot use too obvious means for that.
Technically, we could call the pecking order a "null morality"; like the "null hypothesis" in statistics.
That's forgetting that morality doesn't come from nowhere, it comes from genes too. Because life is full of iterated prisoner’s dilemma, because gene survival requires the survival of your close relatives, because of the way the brain is shaped (like the fact the empathy very likely comes, at least in part, from the way we reuse our own brain circuits to predict the behavior of others).
Moral theories are "artificial constructs", as are all theories. They are generalization, they are abstraction, they can conflict with the "genetic morality", and yes, memes play a huge role in morality. But the core of morality comes from our genes - care for our family, "tit-for-tat with initial cooperation" as the winning strategy for IPD, empathy, ...
Even if ultimately everything comes from the genes, we have to learn some things, while other things come rather automatically.
We educate children to behave nicely to others -- they don't get this ability automatically just because of their genes. On the other hand, children are able to create "Lord of the Flies"-like systems at school without being taught so. Both behaviors are based on evolution, both promote our genes in certain situations, but still one is the default option, and the other must be taught (is transferred by memes).
And by the way, Prisonners' Dilemma is not a perfect model of reality, and the differences are very relevant for this topic. Prisonners' Dilemma or Iterated Prisonners' Dilemma are modelled as series of 1:1 encounters, where the information remains hidden between the interacting players; each player tries to maximize their own utility; and each encounter is scored independently. In real life, people observe what others are doing even when interacting with others; people have families and are willing to sacrifice some of their utility to increase their family's utility; and results of one encounter may influence your survival or death, your health, your prestige etc., which influence the rules of the following encounter. This results in new strategies, such as "signal a membership to a powerful group G, play tit-for-tat with initial cooperation against members of G, and defect against everyone else" which will work if the group G has a majority. Now the problem is how will people agree what is the right group G? In small societies, family can be such group; in larger societies memetic similarity can play the same role -- if you consider that humans are not automatically strategic, why not make a meme M, which teaches them this strategy and at the same times defines group G as "people who share the meme M"? Here comes the morality, religion, football team fans, et cetera.