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RichardKennaway comments on Evolutionary psychology: evolving three eyed monsters - Less Wrong
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Now I'm curious why, as far as I know, no animal has ever evolved an eye in the back of its head. You mention bipedality and descent from trees, but why wouldn't a gazelle or a fly or a small fish get an advantage from one?
Prey animals generally have their eyes on the sides of their heads, which do a good job of covering all directions. And eyes are expensive -- they need a lot of brain.