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pedanterrific comments on Harry Potter and the Methods of Rationality discussion thread, part 15, chapter 84 - Less Wrong
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Presumably they're... well, at the risk of being obvious, the Most Ancient Houses?
Then how could the destruction of one cause their numbers to fall from eight to seven? If it's just a matter of who can trace their roots back the farthest, surely the next-oldest would be bumped up.
Possibly there's some cutoff point, with only houses founded before that point given the Most Ancient label; whether this comes with any official privileges beyond just being old and respected remains to be seen.
It seems like the obvious cut-off point would be the original houses founded when Merlin created the Wizengamot.
By now the Most Ancient label has shifted from being descriptive, to just part of the name