JulianMorrison comments on The Price of Integrity - Less Wrong
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I'd define cowardice as sacrificing a large mind-value to a small but insistent evolutionary-adaptation-value.
Viewed that way, it blends into akrasia. The same problem - insistence.
It's normally defined as a vice of deficiency with respect to concern for risk-taking, when 'cowardice' is used as a trait of character.