The least important audience. An interesting title may draw casual browsers in, but those likely aren't very valuable readers. Most people encountering an academic article will either be looking for it, or will have had it referred to them from some source.

I'm not sure whether that's true. Quite often when doing literature searches I end up with more papers than I have time to read. Going through the citation list of a paper often gives you quickly more papers than you can look at.