ScottL comments on Wrong however unnamed - Less Wrong
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I don't know anything about lichen, but the below is what I assume "more than the sum of" in this context means:
So that means that the complex of f&a acquires different properties, occurs in different conditions, and might - might - have an overall wider 'ecological amplitude'. Which is plausible, but hardly ever proven (I don't recall any direct comparison of rates of reproduction, dissemination, biomass allocation or any other objective measure of population success, not 'body type success', but I have not looked into it.)
This is apples vs. non-apples.