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AspiringRationalist comments on Open Thread May 30 - June 5, 2016 - Less Wrong
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Some people believe that altruism has evolved through helping your relatives or through helping others to help you in return. I was thinking about it; on the surface the idea looks good -- if you already have this system in place, it is easy to see how it benefits those involved -- but that doesn't explain how the system could have appeared in the first place. Anyone knows the standard answer?
Imagine that you are literally the first organism who by random mutation achieved a gene for "helping those who help you". How specifically does this gene increase your fitness, if there is no one else to reciprocate?
Or imagine that you are literally the first organism who by random mutation achieved a gene for "helping your siblings". How specifically does this gene increase your fitness, or the fitness of the gene itself, if your siblings do not have a copy of this gene?
In other words, it seems simple to explain how these kinds of altruism can work when they are already an established system, but it is more difficult to explain how it could work when it is new.
And this all is a huge simplification; for example, I doubt that "helping those who help you" could be achieved by a single mutation, since it involves multiple parts like "noticing that someone helped you", "remembering the individual who helped you" and "helping the individual who helped you in the past". Plus the problem of how to start this chain of mutual cooperation.
My guess is that... nygehvfz pbhyq unir ribyirq guebhtu frkhny fryrpgvba. Yrg'f rkcynva vg ol funevat sbbq jvgu bguref. Svefg, vaqvivqhnyf abgvpr jub vf tbbq ng tngurevat sbbq, naq gurl ribyir nggenpgvba gbjneqf tbbq sbbq pbyyrpgbef. Gung znxrf vzzrqvngr frafr orpnhfr vg vapernfrf fheiviny bs gur puvyqera, vs gurl nyfb trg gur trarf tbbq sbe tngurevat sbbq. Nsgre guvf nggenpgvba rkvfgf jvguva gur fcrpvrf, gur arkg fgrc pbhyq or fvtanyyvat: vs lbh unir fbzr rkgen sbbq lbh qba'g npghnyyl arrq, oevat vg naq ivfvoyl qebc vg arne bgure vaqvivqhnyf, fb gung bguref abgvpr lbh unir zber sbbq guna lbh pna rng. Ntnva, guvf znxrf vzzrqvngr frafr, orpnhfr vg znxrf lbh zber nggenpgvir. Abgvpr ubj arvgure "urycvat lbh eryngvirf" abe "urycvat gubfr jub uryc lbh" jnf arprffnel gb ribyir urycvat vaqvfpevzvangryl. Npghnyyl, gubfr pbhyq unir ribyirq yngre, nf shegure vzcebirzragf bs be nqqvgvbaf gb gur vaqvfpevzvangr urycvat.
If you have a gene that makes you help you siblings, your offspring are reasonably likely to get it too, which benefits their siblings (also your offspring).
I feel like this increases the amount of lucky coincidence needed. Not only I have to randomly get the right mutation, but I also need to have many children (surviving to the age when they can help each other) for reasons completely unrelated to having the mutation. Actually, the mutation may be a bit harmful in the second step, because I may give some of my resources to my siblings instead of my children.
Unfortunately, I am not familiar enough with mathematical models of evolution to evaluate how much this extra burden weighs against your hypothesis.
It seems to me that it doesn't weigh against it very much. A genetic change that causes a not-too-big increase in altruistic behaviour towards likely kin is unlikely to hurt your chances of survival and reproduction a lot.
The first organism with the genetic change doesn't need to be exceptionally well supplied with offspring or anything. (Unless this is an r-selected species for which surviving at all is exceptionally lucky; in that case, it needs to be about as lucky as the bearer of any other not-too-dramatic genetic change has to be.)