dspeyer comments on Harry Potter and the Methods of Rationality discussion thread, part 15, chapter 84 - Less Wrong Discussion
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So, it seems more likely that Quirrel was behind the plot.
The thing about there only being seven houses seems big, though, and as far I can tell isn't from canon. (The list of purebloods, for example, doesn't include Jugson, though 500 years old might not be enough to be Most Ancient. I think we have HPMOR confirmation of Malfoy, Potter, Greengrass, and Longbottom, and I think in canon the only ones that get that description are Malfoy, Black, and maybe Potter (really, Peverell).
The 1926 hint narrows it down to four canon characters (though, of course, Bones might be mistaken). Interestingly enough, all of them were sorted into Slytherin- Tom Riddle, Rosier, Avery, and Lestrange. All of them were Death Eaters, and so it seems most likely it's Tom Riddle. (He would be the last of the female line of the Gaunt family, descended from Salazar Slytherin, which seems like it qualifies for Most Ancient. But I suspect the female line doesn't count for things like the Wizengamot, in canon at least.)
(Interestingly, in canon, Morfin Gaunt was memory-charmed to believe that he was the murderer of Voldemort's parents. Riddle did that to cover up a number of his murders. Even more pieces falling into place.)
Tom Riddle as hero seems... really bizarre, though. Who was Voldemort instead? (It seems implausible that Voldemort could have been an alterego; I suspect quite a bit of his pureblood support came from his lineage.)
Way back in chapter 7, Draco refers to "the Noble and Most Ancient House of Black". That's a fifth noble house.
JKR said Nott was ranked as highly as Malfoy. Doesn't necessarily apply in MOR.
I was going to say the Blacks are all (supposedly) dead in HPMOR at this point, but then I remembered that Sirius is (supposedly) just in Azkaban, not dead yet, and if he's counting the female line (like he would have to for Riddle to count) then there's also Bellatrix.
Maybe Andromeda Black-Tonks can to revive the house even though she was disowned if all the other members are dead or Azkabaned.
There's also Draco, Tonks, and Andromeda. (Andromeda is Tonks' mother, Bellatrix's and Narcissa's sister.) This is all assuming that the female line counts, which it more or less has to.
Yes, I'd assume one of Draco's children would take up the mantle of Black if it came to that.
Or Draco himself.
I assume Draco will remain a Malfoy and that you can't have both.
Can't you? Multiple titles, and even personal union of crowns, are pretty common in RL nobility and royalty. It being allowed would be my default assumption, and I know of no Potterverse evidence to the contrary.
There's a difference between dynasties and titles. Houses are dynasties- lots and lots of people were Habsburgs, even though only one person held a title at any particular time. For example, Philip I was, in 1505, King of Castile, Duke of Burgundy, Duke of Brabant, Duke of Limburg, Duke of Lothier, Duke of Luxemburg, Margrave of Namur, Count Palatine of Burgundy, Count of Artois, Count of Charolais, Count of Flanders, Count of Hainault, Count of Holland, and Count of Zeeland, but only a member of one House- the Habsburgs.
Also, take a look at cadet branches.
It doesn't seem likely that wizards actually have hereditary titles that are linked to locations, since their power comes from magic, not farmland. Lord Malfoy is probably only a Lord because he's on the Wizengamot- Draco doesn't seem to have a courtesy title, as would befit the son of an important position in Muggle Britain.
In MoR, Malfoy is a Noble and Most Ancient House, which presumably comes with a title. In canon, Lucius is neither a Lord, nor on the Wizengamot.
Also, in MoR we have
In that case I'd expect in that case there to be more people whose title is something like "Lord of Malfoy, Black, Longbottom, and Potter".
Oh god, bad fanfiction flashbacks...
Long-form noble titles are used very rarely, because they're so unwieldy, and we've only seen a couple of folks who would be in a position to have multiple titles at all in the sort of detail where the long form would be used. Dumbledore is the only example I know of where they actually used anything longer than a few words. You're right that we might have seen one, but Rowling would likely have found it a bit too complex, and they're not so common that the lack is significant.