In genetics, it seems to be the rule that speculation far outpaces what can actually be known. Countless times I read these papers and they go 'as speculated by X 50 years ago...' (where X is usually Darwin or Fisher). I understand there's some question as to even whether Mendel's peas showed the laws he wanted them to show! Which would indeed exemplify the theory outpacing the practice.
If it's worth saying, but not worth its own post, even in Discussion, it goes here.