Douglas_Knight comments on Stupid Questions November 2015 - Less Wrong Discussion
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It started in X-Files fandom, which is way too late for it to be bang paths. This claims that it started as Action! Mulder, where the ! was specific to the particular modifier, but people don't have a source for that example.
Your link says
and "Canon!Harry" or "HPMOR!Quirrell" is not trait!character, it's world!character which maps to bang paths very well.
Besides, the tendrils of old hacker culture burrow deep into the fabric of memespace... 8-)
Incidentally, the magma computer algebra system uses ! for casting. This is possibly related.
Yes, the fact that the trait usage came first is evidence against the bang-path theory.