I can think of many situations where a zero prior gives rise to tangibly different behavior, and even severe consequences. To take your example, suppose that we (or Omega, since we're going to assume nigh omniscience) asked the person whether JFK was murdered by Lee Harvey Oswald or not, and if they get it wrong, then they are killed/tortured/dust-specked into oblivion/whatever. (let's also assume that the question is clearly defined enough that the person can't play with definitions and just say that God is in everyone and God killed JFK)
However, let me steelman this a bit by somewhat moving the goalposts: if we allow a single random belief to have P=0, then it seems very unlikely that it will have a serious effect. I guess that the above scenario would require that we know that the person has P=0 about something (or have Omega exist), which, if we agree that such a belief will not have much empirical effect, is almost impossible to know. So that's also unlikely.
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Because consciousness is precluded in the thought experiment. The whole idea is that the Zombie World is identical in every way - except it doesn't have this ephemeral consciousness thing.
Therefore the GLUT cannot be conscious, by the very design of the thought experiment it cannot be so. Yet there isn't any logical explanation for the behavior of the zombies without something, somewhere, that is conscious to drive them. That's why the GLUT came into the discussion in the first place - something has to tell the zombies what to do, and that something must be conscious (except it can't be, because the thought experiment precludes it).
Thus, an identical world without consciousness is inconceivable.
So does that mean a GLUT in the zombie world cannot be conscious, but a GLUT in our world (assuming infinite storage space, since apparently we were able to assume that for the zombie world) can be conscious?