Of the examples given, some of them certainly involve controled experiments in the classical sense. Evolutionary biology for example involves tests of genetic drift and speciation in the lab environment. For example, one matter that has been extensively tested in labs is different speciation mechanisms. The founder-effect mechanism is one that is particularly easy to test in a lab. For one major paper on the subject see this paper. A much older example is speciation by hybridization which has been tested in controlled lab environments for about a century now. The oldest I'm aware of in that regard is a 1912 paper by Digby (I haven't read it, and I'd have to go look up the citation but it shouldn't be hard to find ), and there have been many papers since then on the same topic.
Edit:Citation for Digby according to TOA is Digby, L. 1912. The cytology of Primula kewensis and of other related Primula hybrids. Ann. Bot. 26:357-388.
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I think there is definitely potential to the idea, but I don't think you pushed the analogy quite far enough. I can see an analogy between what is presented here and human rights and to Kantian moral philosophy.
Essentially, we can think of human rights as being what many people believe to be an essential bare-minimum conditions on human treatment. I.e. that the class of all "good and just" worlds everybody's human rights will be respected. Here human rights corresponds to the "local rigidity" condition of the subgraph. In general, too, human rights are generally only meaningful for people one immediately interacts with in your social network.
This does simplify the question of just government and moral action in the world (as political philosophers are so desirous of using such arguments). I don't think, however, that the local conditions for human existence are as easy to specify as in the case of a sensor network graph.
In some sense there is a tradition largely inspired by Kant that attempts to do the moral equivalent of what you are talking about: use global regularity conditions (on morals) to describe local conditions (on morals: say the ability to will a moral decision to a universal law). Kant generally just assumed that these local conditions would achieve the necessary global requirements for morality (perhaps this is what he meant by a Kingdom of Ends). For Kant the local conditions on your decision-making were necessary and sufficient conditions for the global moral decision-making.
In your discussion (and in the approach of the paper), however, the local conditions placed (on morals or on each patch) are not sufficient to achieve the global conditions (for morality, or on the embedding). So its a weakening of the approach advanced by Kant. The idea seems to be that once some aspects (but not all) of the local conditions have been worked out one can then piece together the local decision rules into something cohesive.
Edit: I rambled, so I put my other idea into another commend