To be charitable, assume he is aware that pinching a rabbit's leg will trigger afferent nociceptive (pain) neurons, which will carry a signal to the brain, leading to the experience of pain. Your cited research demonstrates this. It does not demonstrate, however, whether the subject has the awareness to reflect upon the factors that contribute to their suffering, such that their reflection can contribute it by further adding negative stimuli, negative stimuli that is generated only by that organism's selfsame reflection.

To be fair, you can't demonstrate this for any human either. That's the problem with consciousness.