Does the CSS make the URLs appear in the middle of text paragraphs, where there was a hyperlink in the online version, or as footnotes or references outside the main text? Nobody really wants the main text to get interrupted by noisy URLs all the time, and on the other hand having URLs written out in the open even in the online version doesn't matter that much if it's in the references section at the end of the document.

The Lynx thing probably isn't good for pleasant reading, but might work for some kind of compromise scheme where you get a document that's very human-readable and can be used for an OCR data restoration that gets you the text and references back, though not most of the formatting.